Arthur Middleton

 

Middleton_Mugshot.pngArthur Middleton completed his Ph.D. during summer 2012. During his time with the Wyoming Cooperative Fish and Wildlife Research Unit, he coordinated the Absaroka Elk Ecology Project in collaboration with the Wyoming Game and Fish Department and the U.S. Fish and Wildlife Service, studying elk migration and elk-wolf interactions in the Absaroka Mountains. The field work on this project is complete, but its final reporting will continue throughout 2012-2013. The primary focus of Arthur's work has been to evaluate the relative influence of predation risk and habitat quality on the nutrition and reproduction of elk, as well as their seasonal migrations. He is also involved in several other projects in Wyoming that investigate the ecology of ungulate migration, antipredator behavior, and reproduction. In September 2012, Arthur began a two-year Donnelly Postdoctoral Fellowship at Yale University. During that time, he will be initiating new research with Emiliano Donadio on the ecology of a high Andean food web in Argentina, where pumas and condors interact strongly with native camelids. Before coming to Wyoming, Arthur received a master’s degree in environmental management from Yale and a bachelor’s degree in English and government from Bowdoin College. Most of his prior experience was with birds of prey - as a falconer, he trained raptors species for public education and hunting, and as a field biologist, he worked with swallow-tailed kites in South Carolina, harpy eagles in Panama, and bald eagles in New York. Arthur is the recipient of a Thomas J. Watson Fellowship, a Doris Duke Conservation Fellowship, and a Resident Fellowship at the Buffalo Bill Historical Center in Cody, Wyoming.

Arthur defended his dissertation in July, 2012. You can watch his public seminar via WyoCast.

Download Arthur's CV


Publications

Middleton, A.D., M.J. Kauffman, D.E. McWhirter, M.D. Jimenez, J.G. Cook, R.C. Cook, S.E. Albeke, H. Sawyer, and P.J. White. 2013. Linking antipredator behavior to prey demography reveals limited risk effects of an actively hunting large carnivore. Ecology Letters 16:1023-1030. Original article, ScienceNOW, Casper Star-Tribune, Jackson Hole News and Guide, Missoulian.

Middleton, A.D., M.J. Kauffman, D.E. McWhirter, J.G. Cook, R.C. Cook, A.A. Nelson, M.D. Jimenez, and R.W. Klaver. 2013. Rejoinder: Challenge and opportunity in the study of ungulate migration amid environmental change. Ecology 94:1280-86.

Middleton, A.D., M.J. Kauffman, D.E. McWhirter, J.G. Cook, R.C. Cook, A.A. Nelson, M.D. Jimenez, and R.W. Klaver. 2013. Animal migration amid shifting patterns of phenology and predation: Lessons from a Yellowstone elk herd. Ecology 94:1245-56. Original article, Connecticut Public Radio (audio)Wyoming Public Radio's Open Spaces (audio), Billings Gazette, Casper Star-Tribune, Jackson Hole News and GuideSan Francisco Chronicle

Middleton, A.D., T.A. Morrison, J.K. Fortin, M.J. Kauffman, C.T. Robbins, K.M. Proffitt, P.J. White, D.E. McWhirter, T.M. Koel, D. Brimeyer, and W.S. Fairbanks. 2013. Grizzly bear predation links the loss of native trout to the demography of migratory elk in Yellowstone. Proceedings of the Royal Society B: Biological Sciences 280: 20130870. Original articleNature Research Highlight, ScienceNOWBillings Gazette, Casper Star-Tribune, Jackson Hole News and Guide, Bozeman Chronicle, Conservation Magazine, Agence France-PresseLe MondeSpiegel

Christianson, D., R.W. Klaver, A.D. Middleton, and M.J. Kauffman. 2013. Confounded winter and spring phenoclimatology on large herbivore ranges. Landscape Ecology 28:427-437.

Sawyer, H., M.J. Kauffman, A.D. Middleton, T.A. Morrison, R. Nielsen, T. Wyckoff. 2013. A framework for understanding semi-permeable barrier effects on migratory ungulates. Journal of Applied Ecology 50:68-78.

Nelson, A.A., M.J. Kauffman, A.D. Middleton, M.D. Jimenez, D.E. McWhirter, J. Barber, and K. Gerow. 2012. Elk migration patterns and human activity influence wolf habitat use in the Greater Yellowstone Ecosystem. Ecological Applications 22:2293-2307.pdf.jpg

Cross, P.C., E.K. Cole, A.P. Dobson, W.H. Edwards, K.L. Hamlin, G. Luikart, A.D. Middleton, B.M. Scurlock, and P.J. White. 2010. Probable causes of increasing elk brucellosis in the Greater Yellowstone Ecosystem. Ecological Applications 20: 278-288. pdf.jpg

Martínez del Rio, C. and A.D. Middleton. 2010. Laws for ecology? (book review). Ecology 91:1244-1245. pdf.jpg

Pauli, J.N., J.P. Whiteman, M. Riley, and A.D. Middleton. 2010. Defining noninvasive for sampling of vertebrates. Conservation Biology 24:349-352. pdf.jpg

In review/revision

Koch, B.J., A.D. Middleton, R.O. Hall, and B.L. Peckarsky. In revision. Secondary production integrates behavioral, physiological, and life history effects in diverse prey taxa.

In preparation

Middleton, A.D., M.J. Kauffman, and C. Martinez del Rio. Prey nutritional condition: a common influence on antipredator behavior and the 'ecology of fear' (Synthesis).

Nelson, A.A., M.J. Kauffman, A.D. Middleton, M.D. Jimenez, and D.E. McWhirter, and K. Gerow. A comparison of wolf depredation sites in areas with migratory and resident prey.


Selected presentations

Middleton, A.D., M.J. Kauffman, M.D. Jimenez, D.E. McWhirter, A.A. Nelson, J.C. Cook, and R.C. Cook. Evaluating a proposed non-consumptive effect of wolves on elk in Yellowstone. Annual Meeting of The Wildlife Society, Kona, Hawaii, November 2011 (and Annual Meeting of the Wyoming Chapter of The Wildlife Society, Jackson, WY, December 2011).

Middleton, A.D. Animal migration amid shifting patterns of phenology and predation: Lessons from a Yellowstone elk herd. Invited seminar, Department of Forest and Wildlife Ecology, University of Wisconsin - Madison, Madison, WI,  23 September 2011.

Middleton, A.D. Changing times in Wyoming elk country: Large carnivores, drought, and elk migration. Rocky Mountain Elk Foundation, Annual Habitat Council Retreat, Cody, WY, 11 June 2011.

Middleton, A.D. The ecology of elk migration in the Greater Yellowstone Ecosystem. Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata, Argentina, 18 March 2011.pdf.jpg

Middleton, A.D., M.J. Kauffman, D.E. McWhirter, M.D. Jimenez,  J.G. Cook, R.C. Cook, and A.A. Nelson. Do predators influence the condition and reproduction of their prey? Annual Meeting of the Wyoming Chapter of The Wildlife Society. Lander, WY, November 2010.

Middleton, A.D., M.J. Kauffman, D.E. McWhirter, J.G. Cook, R.C. Cook, A.A. Nelson, M.D. Jimenez, and R.W. Klaver. Large carnivore recovery and long-term drought reduce the benefits of migration for a Yellowstone elk herd. Invited seminar, Faculty Seminar Series, Yale School of Forestry and Environmental Studies, New Haven, CT, October 2010. 

Middleton, A.D., M.J. Kauffman, D.E. McWhirter, J.G. Cook, R.C. Cook, A.A. Nelson, M.D. Jimenez, and R.W. Klaver. Large carnivore recovery and long-term drought reduce the benefits of migration for a Yellowstone elk herd. Annual Meeting of the Ecological Society of America, Pittsburgh, PA, August 2010.

Middleton, A.D., M.J. Kauffman, D.E. McWhirter, J.G. Cook, R.C. Cook, A.A. Nelson, M.D. Jimenez, and R.W. Klaver. Large carnivore recovery and severe drought reduce the benefits of migration for a Yellowstone elk herd. Annual Meeting of the American Society of Mammalogists, Laramie, WY, June 2010. Reported by Science News. 

Middleton, A.D. Drought, predation, and migratory Clarks Fork elk. Several talks given at the Wyoming Game and Fish Department's Cody All-Region Meeting and Terrestrial Habitat Section Annual Meeting and the U.S. Fish and Wildlife Service Wolf Recovery Annual Meeting. Cody and Kelly, WY and West Yellowstone, MT. May-June 2010.

Middleton, A.D. Drought, predation, and migratory Clarks Fork elk. Invited public presentations at the Buffalo Bill Historical Center in Cody, WY and Teton County Library in Jackson, WY. May 2010. Reported by the Billings Gazette.

Middleton, A.D. The shifting benefits of migration for Clarks Fork elk. Annual Meeting of the Wyoming Chapter of The Wildlife Society. Cody, WY, November 2009.


Scholarships and Fellowships 

2011                   Scott-Walter Funds, Department of Zoology & Physiology, University of Wyoming
2011  Dick & Lynne Cheney International Funds, University of Wyoming
2010  L. Floyd Clarke Graduate Scholar Award, University of Wyoming
2009-10 Resident Fellow, Cody Institute for Western Studies, Buffalo Bill Historical Center
2008       NSF Graduate Research Fellowship - Honorable Mention 
2008 Plummer Scholarship, Environment & Natural Resources, University of Wyoming
2007-08 NSF-EPSCoR Fellowship, Program in Ecology, University of Wyoming
2007 Paul D. Camp Award, Yale University
2006 Gilman Ordway Environmental Scholarship, Yale University
2006-07 Doris Duke Conservation Fellowship, Yale University
2003-04 Thomas J. Watson Fellowship, Thomas J. Watson Foundation
   

Projects

Absaroka Elk Ecology Project

Absaroka Wolf-Livestock Project